The Dude wrote:
I would rebut the argument by pointing out that "survival of the fittest" is a very poor summation of the theory of evolution that was only borrowed by Darwin in the fifth edition of Origin of the Species. It is such a dumbed-down metaphor that it isn't even worth defending. Ridicule it all you want, if that's the very best you can do.
He's arguing that the concept of "fitness" is a tautology. That is more central to evolutionary biology now. The term owes its origin to a saying that you correctly state is oversimplified, but that doesn't mean the concept of a fitness landscape isn't worth defending. Will is arguing that the concept of natural selection a tautology because saying organisms who are more fit in a given environment are more likely to propagate through time is a tautology. In his mind "fit" just means, "those that propagate through time." That's not true, and rather obviously so if you are armed with a real basic understanding of the idea, but that's where he's coming from.